Aegean Bryophyte Atlas

The distribution of bryophytes known from the Chalkidiki peninsulas and Aegean islands, Greece.
An atlas project.

compiled by Burkhard Biel,
generously supported by Tom Blockeel

Based on the work of the late Ruprecht Düll (2014) in his book
”A Survey of the Bryophytes known from the Aegean Islands“, this atlas project shows the distribution of the bryophytes of the Aegean area on maps within the boundaries of Düll's work (he died on 07.06.2014, shortly before its publication).
The main aim of this project is to present quick and up-to-date results from surveys of bryophytes in the Aegean area, with the help of distribution maps, thus completing and continuing Düll's work. It should encourage further exploration of the Greek bryophyte flora, which is much needed.
The data from Düll's work have been critically evaluated by Tom Blockeel and provided with comments if necessary. Additional data, both published and unpublished, have been included (see below).

Most of the reports included in this work have only been assessed for their credibility; there is no guarantee of their correctness.

Up to now, data are included from the three peninsulas of Chalkidiki (Athos, Kassandra, Sithonia) and the following 31 islands: Alonnisos, Amorgos, Andiparos, Chios, Crete, Delos, Egina, Euboea, Fourni, Gavdopoula, Ikaria, Kalymnos, Karpathos, Kos, Kythera, Kythnos, Lesvos, Methana, Paros, Patmos, Rhodos, Samothrace, Samos, Santorini (Thira, Thirasia, Nea Kaimeni), Siphnos, Serifos, Sikinos, Skiathos, Skopelos, Thasos, Tilos. 
The location of these peninsulas and islands in the Aegean Sea is shown on a separate map.

This project will be updated continuously, as soon as new data are available.

Everyone who is interested in the Greek bryophytes is invited to support this project with their data. The aim is to compile and present as complete a picture of their distribution as possible. The contact address is .

I am very grateful to Beáta Papp who promised her support for this project from the first inquiry onwards and offered the ECCB website with the server of the Hungarian Natural History Museum as an Internet platform. Many thanks too to Erzsébet Szurdoki who constructed this website.

Burkhard Biel


The following publications have been evaluated:
Blockeel, T.L. 2012. Notes on the Bryophytes of the schistose valleys of western Crete, including Asterella africana new to Greece, and an assessment of the conservation status of Rhamphidium purpuratum and Jungermannia handelii. - Journal of Bryology 34: 292-296.
Blockeel, T.L. & Nieuwkoop, J.A. 2016. The bryophyte flora of Lesbos, Greece. - Herzogia 29: 1-34.
Blockeel, T.L. 2016. The mosses Aschisma carniolicumCampylostelium strictum and Timmiella flexiseta on the Eastern Aegean Islands, new to Greece, and a second locality for Syntrichia latifolia. - Herzogia 29: 52-58.

Düll, R. 2014. A survey of the Bryophytes known from the Aegean islands. - Jena: Weissdorn-Verlag.
Kay, Q. 1969. Some mosses from Khios. - Revue Bryologique Lichénologique 36: 625-627.
Papp, B. (2007): Reports 1-7. In: Natcheva, R., Tsakiri, E., Dihoru, G. (eds.): New bryophyte records in the Balkans: 1. – Phytologica Balcanica 13(1): 102.

Unpublished data have been provided by
B. Papp: Crete (1996, 1997, 2001), Halkidiki (Kassandra) (2003), Rhodos (2006)
H. Sipman: Andiparos (1998), Kalymnos (2000), Kos (2000), Paros (1998), Santorini (1990, 1998, 2000)
B. Biel: Kythnos-Serifos-Siphnos (2003-4), Alonnisos-Skiathos-Skopelos (2005), Ikaria (2005), Sikinos (2005), Samothrace (2006-13), Amorgos (2015)

All known species and subspecies (including the most important synonyms) of the bryophytes from the Aegean islands are presented in two lists, one for the liverworts and one for the mosses, in alphabetical order. A click on a species name opens a separate window with the relevant distribution map.
The maps show the Greek Aegean area divided into phytogeographical regions (according to Flora Hellenica, Strid & Tan 1977) abbreviated as follows: EAe = East Aegean, EC = East Central, Kik = Kiklades, KK = Kriti-Karpathos, NAe = North Aegean, NE = North East, Pe = Peloponnese, StE = Sterea Ellas, WAe = West Aegean.
The records are shown on the maps with one dot for each island or peninsula (except Kriti: four dots)
 with different symbols for three time periods as  white dot <1950 <  pink dot  <1990< red dot current status.

Appropriate comments are added at the top right side of the maps.

Last updated: 30.11.2016

The list of liverworts (Anthocerotae, Hepaticae) currently contains 122 taxa, including 13 synonyms (italic font).

The list of mosses (Bryatae) currently contains 439 taxa, including 92 synonyms (italic font).